L. Salgadoa, P.A. Gallina and A. Paulina Carabajal

The titanosaur sauropod Bonatitan reigi is redescribed. The material collected, originally interpreted as pertaining to two different individuals, is reorganised in five individuals, and the original type specimen is separated into three different individuals. One of the braincases is designated as a new type specimen. Some materials are described by the first time (sacral ribs, distal caudal, chevrons, metacarpals, astragalus and metatarsals), others reinterpreted as different bones (‘ulna’ and ‘radius’). The diagnosis of B. reigi is emended, removing some of the original characters (longitudinal groove located on the suture between the parietals that continues posteriorly over the supraoccipital to the foramen magnum) and adding some new (small paired pits on the frontals and posterior ridge of the metacarpal IV). The phylogenetic analysis does not support B. reigi as a member of the Saltasaurinae, but rather as a basal member of a broad clade of sauropods that in turn is recovered as the sister group of the Saltasauridae.

L. Salgado, P.A. Gallina & A. Paulina Carabajal (2014): Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina), Historical Biology: An International Journal of Paleobiology, DOI: 10.1080/08912963.2014.894038


 

Jonathas S. Bittencourt, Andrea B. Arcucci, Claudia A. Marsicano and Max C. Langer


Lewisuchus admixtus is an enigmatic early dinosauriform from the Chañares Formation, Ladinian of Argentina, which has been recently considered a member of Silesauridae. Yet, it differs markedly from Late Triassic silesaurids in dental and vertebral anatomy. Indeed, a detailed redescription of its holotype allowed the identification of several features of the skeleton previously unrecognized amongst silesaurids. These include pterygoid teeth, a dorsomedial posttemporal opening on the otoccipital, foramina associated with cranial nerves X–XII on the caudal region of the prootic–otoccipital, and postaxial neck/trunk vertebrae with craniocaudally expanded neural spines. The presence of a single row of presacral scutes was also confirmed. Some elements previously referred to, or found associated with, the holotype, including a lower jaw, pedal elements and an astragalus, more probably correspond to proterochampsid remains. The anatomical information available for the holotype of L. admixtus was rescored into a new phylogenetic dataset for dinosauromorphs, mostly based on previous works. Lewisuchus admixtus and Pseudolagosuchus major are treated as distinct OTUs because their preserved skeletons mostly lack overlapping parts. The parsimony analysis supports the basal position of L. admixtus within dinosauriforms, prior to the silesaurid–dinosaur split, rather than at the base of Silesauridae. This suggests that a higher number of early dinosauriform clades branched in the Middle and Late Triassic than previously suggested.

 
Jonathas S. Bittencourt, Andrea B. Arcucci, Claudia A. Marsicano & Max C. Langer (2014): Osteology of the Middle Triassic archosaur Lewisuchus admixtus Romer (Chañares Formation, Argentina), its inclusivity, and relationships amongst early dinosauromorphs, Journal of Systematic Palaeontology, DOI: 10.1080/14772019.2013.878758


 
Ariel H. Méndez
The caudal vertebrae of abelisaurid theropods show notable variations in the shape of the transverse process, among other characters, and these distinguish the two subclases Brachyrostra and Carnotaurini. The vertebrae in the proximal third of the tail in abelisaurids, especially derived taxa, show adaptations that provide considerable rigidity, when compared to other theropod dinosaurs, especially Maniraptora, which have highly flexible tails. This continues the considerable rigidity in the neck, trunk, and sacral region in abelisaurids, and is consistent with the hypothesis that South American abelisaurids evolved in isolation from those that inhabited India and Madagascar.
 Méndez, A.H. 2014. The caudal vertebral series in abelisaurid dinosaurs. Acta Palaeontologica Polonica 59 (1): 99–107.


Mariela S. Fernández and Ashu Khosla

The eggshell oospecies from India and Argentina are compared and reviewed in detail. These eggshells resemble each other in having a nodular outer surface ornamentation and clearly arched growth lines of the shell units. Microstructurally, the eggshell oospecies belonging to the oofamily Megaloolithidae shows fan-like shell units, which are sharply separated from each other throughout the thickness of the eggshell and can be traced up to the surface of the eggshell. Comparisons between four oospecies from India and Argentina reveal three groupings, which show similarities between megaloolithids of both countries: (1) Megaloolithus jabalpurensis, M. matleyi and M. patagonicus; (2) M. cylindricus, M. rahioliensis and Tipo 1d; and (3) M. megadermus and Tipo 1e. The other two types of eggshell oospecies from India and Argentina show partially fused external nodes and shell units. As a result, growth lines enter into the adjacent shell units with a marked concavity. A new oogenus Fusioolithus have been erected due to fusion between shell units and tubospherulitic morphotype, which include two new oospecies F. baghensis and F. berthei. Till date, morphostructurally, a total of 15 eggshell oospecies belonging to different oofamilies have been recorded from India and seven oospecies from Argentina.

Mariela S. Fernández & Ashu Khosla, Historical Biology (2014): Parataxonomic review of the Upper Cretaceous dinosaur eggshells belonging to the oofamily Megaloolithidae from India and Argentina, Historical Biology: An International Journal of Paleobiology, DOI: 10.1080/08912963.2013.871718.

 
 
 


Bernardo González Riga y Leonardo Ortíz David

Quetecsaurus rusconii gen. et sp. nov. is a new titanosaur (Dinosauria, Sauropoda) from the Neuquén Basin of Mendoza Province, Argentina. The specimen comes from red mudstones of the Cerro Lisandro Formation (middle-late Turonian, Upper Cretaceous), and is the first sauropod with well-preserved remains to be discovered in this formation. The holotype includes a postorbital, teeth, the atlas, a posterior cervical vertebra, an incomplete dorsal vertebra, a posterior caudal centrum, dorsal ribs, a coracoid, fragments of a humerus, radius and ulna, and five metacarpals. It is characterized by the following combination of autapomorphies: (1) intercentrum of atlas with a prominent anteroventral border and expanded posteroventral processes; (2) posterior cervical neural spines with incipient lateral expansions; and (3) humerus with strongly sigmoid proximal border, rounded proximomedial border, and angular proximolateral corner. A preliminary cladistic analysis placed Quetecsaurus as a sister taxon of Lognkosauria (Mendozasaurus + Futalognkosaurus). Quetecsaurus shares with the lognkosaurs the presence of cervical neural spines with ‘lateral laminae’, but relatively reduced in comparison with those taxa. This discovery provides new information on the diagnosis of Lognkosauria within South American titanosaurs.

González Riga, B. y Ortíz David, L. 2014.A New titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous (Cerro Lisandro Formation) of Mendoza Province, Argentina. Ameghiniana 51(1):3-25.

 

Lucas Ernesto Fiorelli, Gerald Grellet-Tinner, Eloisa Argañaraz y Leonardo Salgado

El sitio de nidificación de Sanagasta (Cretácico), La Rioja, Argentina, resolvió aspectos sobre la reproducción de los neosaurópodos, evidenciando comportamientos coloniales, filopatría, y su reproducción oportunista dependiente de su paleoambiente termal. Aquí describimos la tafonomía del sitio determinando sus patrones bioestratinómicos y diagenéticos. El agrupamiento oológico 3D está bien preservado, sugiriendo asociaciones autóctonas representadas por oviposiciones in situ, sin estructura de nido. El nivel portador presenta una biofábrica matriz-soportada y una clasificación genética de acumulación biogénica “intrínseca”; esto implica comportamiento gregario y colonización preferencial del sitio. La biocenosis corresponde a la producción biogénica (huevos), con la cáscara diagenizada por hidrotermalismo y microorganismos extremófilos. La tanatocenosis y el tiempo promedio fueron nulos por el inmediato enterramiento de los huevos luego de la oviposición indicando que no existió una etapa bioestratinómica. La tafocenosis representa el carácter tafonómico más significativo. Las cáscaras presentan alteraciones por soluciones hidrotermales, mientras la recalcificación y silicificación representan las mineralizaciones más frecuentes. La presencia de microchert, calcedonia, microesferas de ópalo-A, agregados de epidoto y minerales de arcilla reemplazando sectores de la cáscara y las membranas del huevo, confirman una alteración hidrotermal bajo condiciones excepcionales. La presencia de microorganismos fósiles (estromatolitos, cianobacterias y diatomeas), numerosas macro-estructuras, microfacies y microfábricas hidrotermales, y los atributos tafonómicos oológicos, sugiere que el sitio de nidificación de Sanagasta representa un particular ejemplo de tafocenosis y un Lagerstätte. El sitio de nidificación de neosaurópodos de Sanagasta es el primer caso fósil documentado sobre la reproducción de un grupo de dinosaurios dependiente de su ambiente.

Fiorelli, L. E., Grellet-Tinner, G., Argañaraz, E. y Salgado, L. 2013. Tafonomía del sitio de nidificación de Neosaurópodos de Sanagasta (La Rioja, Argentina): Ejemplo de preservación excepcional en un paleoambiente hidrotermal del Cretácico. Ameghiniana 50 (4): 389 – 406.

 

Cretaceous Small Scavengers

 
Silvina de Valais, Sebastián Apesteguía, Alberto C. Garrido
Ecological relationships among fossil vertebrate groups are interpreted based on evidence of modification features and paleopathologies on fossil bones. Here we describe an ichnological assemblage composed of trace fossils on reptile bones, mainly sphenodontids, crocodyliforms and maniraptoran theropods. They all come from La Buitrera, an early Late Cretaceous locality in the Candeleros Formation of northwestern Patagonia, Argentina. This locality is significant because of the abundance of small to medium-sized vertebrates. The abundant ichnological record includes traces on bones, most of them attributable to tetrapods. These latter traces include tooth marks that provide evidence of feeding activities made during the sub-aerial exposure of tetrapod carcasses. Other traces are attributable to arthropods or roots. The totality of evidence provides an uncommon insight into paleoecological aspects of a Late Cretaceous southern ecosystem.
de Valais S, Apesteguía S, Garrido AC (2012) Cretaceous Small Scavengers: Feeding Traces in Tetrapod Bones from Patagonia, Argentina. PLoS ONE 7(1): e29841. doi:10.1371/journal.pone.0029841

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